01/2003-07/2007

Ph.D. candidate, Molecular Cellular Biology, Faculty of Medicine, Laval University, Québec, Canada (Dr. Robert Viger) 31/07/2007 Ph.D.

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07/2007-03/2009

Postdoctoral fellow, Ontology and Reproduction Unit CHUQ Research Centre, Laval University, Québec, Canada (Dr. Robert Viger)

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04/2009-04/2010

Postdoctoral fellow, Goodman Cancer Centre,　Faculty of Medicine, McGill University, Montreal, Canada (Dr. Yang XJ)

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05/2010-03/2012

Young Research Scientist, Disease Mechanism Core, RIKEN Brain Science Institute, Saitama, Japan (Dr. Adrian Moore)_My work was introduced by RIKEN NEWS (p14)

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04/2012-08/2014

Assistant Professor, Laboratory for Genetic Code, Graduate School of Life and Medical Sciences, Doshisha University,  Kyoto, Japan_My work was introduced by Doshisha University News Letter LIAISON (p7)